Levitt in 1974, proposed a synthetic theory on the mechanism of stomatal movement, taking all earlier observations into account. The observed facts are:
(a) During opening of stoma, starch disappears from the guard cells.
(b) Photoactive opening is dependent on the chloroplast of the guard cells.
(c) Low CO2 concentration causes stomatal opening.
(d) High pH in guard cells cause stomatal opening.
(e) There is a direct relation between K+ accumulation and stomatal opening.