Consider the following characteristics of herbivore digestive systems and habits.
1. Cows have 4 chambered stomachs and chew cud.
2. Rabbits eat their droppings called cecotropes.
3. Pandas have short digestive system and a small presence of bacteria in the gut.
4. Elephants have a large cecum (the part connecting the two intestines) containing fermenting bacteria.
Which of the above characteristics help food to remain longer in the digestive tract?
1, 2, 4