Discuss the effect of various environmental and plant factors on the process of photosynthesis.
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Solution
Photosynthesis
is affected by both internal (plant) and external (environmental) factors.
The
plant factors include the number, size, age and orientation of leaves, mesophyll
cells and chloroplasts, internal O2 concentration and amount of
chlorophyll. The plant factors are dependent on the genetic predisposition and
the growth of the plant.
The environmental factors include the availability of
sunlight, temperature, CO2 concentration and water.
Light: It is an
essential factor for photosynthesis. It affects the rate of photosynthesis as:
Light
intensity: There is a linear relationship between incident light and CO2
fixation at low light intensities. At higher light intensities, gradually the
rate does not show further increase as other factors become limiting. The light
saturation occurs at 10 percent of the total sunlight available to plants.
Increase in incident light beyond a point causes the breakdown of chlorphyll
and thus resulting in decrease in photosynthesis. Hence, except for plants in
shade or in dense forests, light rarely become a limiting factor.
Light quality:
Light between 400-700 nm wavelengths constitute the photosynthetically active
radiation (PAR). Maximum photosynthesis takes place in red and blue light of
the visible spectrum and minimum photosynthesis takes place in green light.
Duration
of light: Light duration does not affect the rate of photosynthesis, but it
affects the overall photosynthesis.
Carbon Dioxide Concentration: It is a major
limiting factor influencing the rate of photosynthesis. The concentration of
CO2 is very low in the atmosphere which is far below the requirement for
optimum photosynthesis. Increase in concentration up to 0.05 percent can cause
an increase in the rate of photosynthesis. But beyond this level, it becomes
damaging over longer periods.
Temperature: Photosynthesis can take place over a
wide range of temperatures. The light reactions are temperature sensitive, but
they are affected to a much lesser extent. The dark reactions being enzymatic
are temperature controlled. Again, the temperature optimum for photosynthesis
also depends on the habitat the plants are adapted to. The C4 plants
respond to higher temperatures and they show higher rate of photosynthesis,
while C3 plants have much lower temperature optimum.
Water: Water is one
of the raw materials utilized for the process of photosynthesis. Photosynthetic
process utilizes less than 1% of the water absorbed by a plant; hence it is
rarely a limiting factor in photosynthesis. Water stress causes the stomata to
close, hence reducing the CO2 availability as gaseous exchange could not
occur. Also, water stress makes leaves wilt, thus reducing the surface area of
the leaves and the metabolic activity reduces as well. Thus, the effect of
water as a factor is more through its effect on the plant, rather than directly
on photosynthesis.