The correct option is
A High temperature-
+, Lack of oxygen-
+, Moist environment-
+, Lignin and chitin in detritus-
−Decomposers and transformers are the living components of the ecosystem and they are fungi and bacteria. Decomposers attack the dead remains of producers and consumers and degrade the complex organic substances into simpler compounds. The simple organic matters are then attacked by another kind of bacteria, and transformers which change these organic compounds into the inorganic forms that are suitable for reuse by producers or green plants.
The factors affecting microbial decomposition: Temperature is an important factor in the biological of a compost heap. Low outside temperatures during the winter months slow the decomposition process, while warmer temperatures speed it up. During the warmer months of the year, intense microbial activity inside the heap caused composting to proceed at extremely high temperatures. The high temperatures are beneficial to the gardener because they kill weed seeds and germs that could be detrimental to vegetation. The next holds at 100 °F for a while and different microbes predominate. Then finally, the ambient phase where the pleasant earthly odor originates and material has produced compost.
Lack of oxygen: Composting can be defined in the terms of availability of oxygen. Aerobic decomposition means that the active microbes in the heap require oxygen, while in anaerobic decomposition, the active microbes do not require oxygen to live and grow. Temperature, moisture content, the size of bacterial populations, and availability of nutrients limit and determine how much oxygen your heap uses.
Moist environment: The amount of moisture in your heap should be as high as possible, while still allowing air to filter into the pore spaces for the benefit of aerobic bacteria. Individual bacterial hold various percentages of moisture in compost and determine the amount of water that can be added.
lignin and chitin in detritus: Even, within same climatic conditions decomposition rate becomes slower, if the detritus is rich in lignin and chitin. The nitrogen-rich detritus, on the other hand, having low amounts of lignin, decomposes relatively rapidly.
So the correct option is 'high temperature +, lack of oxygen +, moist environment +, lignin and chitin in detritus-'.