In maize, coloured endosperm (C) is dominant over colourless (c) and full endosperm (R) is dominant over shrunken (r), When a dihybrid of F1 generation was test crossed it produced four phenotypes in the following percentage: Coloured-Full = 45%, Coloured-Shrunken = 5%, Colourless-Full = 4% Colourless-Shrunken = 46%