Match the following.
Column - I Column - II
(i). Epidermis (a). Nutritive layer
(ii). Endothecium (b). Dehiscence of anther
(iii). Tapetum (c). Protective layer
(iv). Middle layers (d). Short-lived layer
(i) - (c), (ii) - (b), (iii) - (a), (iv) - (d)
Anther wall consists of four layers- epidermis, endothecium, middle layers and tapetum. The epidermis is thick-walled and gives protection. Endothecial cells have fibrous thickenings which are hygroscopic helping in anther dehiscence. Middle layers are thin walled and short-lived while tapetum is the innermost layer that supplies nutrients to developing pollen.