The correct option is D All of the above
Various abiotic factors associated with damage to food grains are:
1. Temperature: If food is stored at lower temperature, then enzymes, insects and the other micro-organisms, which spoil the food, become less active. The damage to the stored food is decreased.
2. Moisture: For safe storage of food grains, it is essential that the moisture in the food grains should not be more than 14% by weight.
3. Humidity: Growth of moulds and fungi in food grains takes place due to humidity of air.
4. Material of the container: Food grains should not be stored in lead container, because lead is a toxic metal.