Apertures are areas on the walls of a pollen grain, where the wall is thinner and/or softer. For germination it is necessary that the pollen tube can reach out from the inside of the pollen grain and transport the sperm to the egg deep down in the pistil. The apertures are the places where the pollen tube is able to break through the (elsewhere very tough) pollen wall.
The number and configuration of apertures are often very exactly characteristic of different groups of plants. The biggest class of plant species, the Eudicots, usually have three apertures in each pollen grain.
ADDITIONAL KNOWLEDGE
In optics, an aperture is a hole or an opening through which light travels. More specifically, the aperture and focal length of an optical system determine the cone angle of a bundle of rays that come to a focus in the image plane. The aperture determines how collimated the admitted rays are, which is of great importance for the appearance at the image plane.[2] If an aperture is narrow, then highly collimated rays are admitted, resulting in a sharp focus at the image plane. A wide aperture admits uncollimated rays, resulting in a sharp focus only for rays coming from a certain distance. This means that a wide aperture results in an image that is sharp for things at the correct distance. The aperture also determines how many of the incoming rays are actually admitted and thus how much light reaches the image plane (the narrower the aperture, the darker the image for a given exposure time). In the human eye, the pupil is the aperture.