S1 gives the definition of Palaeobotany. There is a clear sequence of Q followed by R as R gives an example of the evolution mentioned in Q. Similarly, there is a clear sequence of P followed by S as there is a link of fossils between them. Now, the sequence QRPS makes more sense than PSQR as there is no connection between statements S and Q. On the other hand, S is a good precursor to S6 as S6 mentions things like climate, temperature and topography that are closely related to the geologist's area of investigation.
So the answer is QRPS.