A lenticel is a porous tissue consisting of cells with large intercellular spaces in the periderm of the secondarily thickened organs and the bark of woody stems and roots of dicotyledonous flowering plants. It functions as a pore, providing a pathway for the direct exchange of gases between the internal tissues and atmosphere through the bark, which is otherwise impermeable to gases. The shape of lenticels is one of the characteristics used for tree identification. In plant bodies that produce secondary growth, lenticels promote gas exchange of oxygen, carbon dioxide, and water vapor. Lenticel formation usually begins beneath stomatal complexes during primary growth preceding the development of the first periderm.