Loss of grains during storage can occur because of following factors:
1. Biotic factors: During storage, insects, fungi, mites and rodents may grow. The growth of these organisms can lead to grain spoilage, which eventually leads to loss of grains.
2. Abiotic factors: Certain abiotic factors such as temperature and moisture content can promote the growth of microorganisms. This leads to grain spoilage as well as grain loss.