A biologist researched the rate of evolutionary change in a butterfly population on a group of islands over a period of several decades.
He determined that the average length of the proboscis (external mouthpart) of butterflies in a particular population decreased dramatically during a ten year period in which annual rainfall was reduced by 60%.
During that same ten year period, the biologist observed a reduction in the average size of flowers produced by the indigenous plants in the area. In the five years after annual rainfall returned to normal levels, the biologist discovered that the average length of the proboscis increased to the original lengths observed. This reversal in proboscis length can best be explained by which of the following?