By cross pollinating certain tobacco species, some fertile hybrids were obtained by
In tobacco, due to the continuous self-pollination, homozygosity increases. It results in the occurrence of inbreeding depression, and the plants are called Inbred lines. When these inbred lines are subjected to hybridization, fertile hybrids are obtained. This is called Hybrid vigour or heterosis. This was developed by Kolreuter.
So, the correct option is ‘Kolreuter’.