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Question

Differentiate Between.
C3 and C4 cycles.

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Solution

In C3 cycle (also known as Calvin cycle), CO2 is fixed when it is accepted by a 5-C compound RuBP to form two molecules of a 3-C compound, 3- osphoglyceric acid, the first product of photosynthesis. Ultimately RuBP is regenerated. On the other hand, in C4 cycle (also known asHatch and Slack cycle), the first compound in photosynthesis is a 4-C compound, the oxaloacetate. The chloroplasts of mesophyll cells of C4 plants though have typical granal stacking and other usual features of a chloroplast from a green plant, they lack ribulose bisphosphate carboxylase activity which is present in the chloroplasts of mesophyll cells of C3 plants: The C4 cycle is comparatively more energy-expensive than the C3 cycle, as in C3 plants, there is an approximate requirement of 3 ATP and 2 NADPH per molecule of CO2 assimilated, whereas in C4 plants, additional ATPs are required for conversion of pyruvate to phosphoenol pyruvate of decarboxylation of oxaloacetate to phosphoenol pyruvate. In general, C4 pathway of photosynthesis in tropical habitats is a device to reduce water loss through transpiration by means of partial stomatal closure so that the plants lose less water for CO2 fixation compared to C3 plants and can make more economic use of water for growth.

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