Read the passage and answer the following question.
A total of 12 bird species having distinct diets, behavior, coloration and beak shapes inhabit a large volcanic island in the South Atlantic 500 miles from the Coast of Africa not found on the mainland. Two species are flightless herbivores. A bird found on the mainland is related to these bird species but it does not fly over water but can occasionally survive at sea in a storm. There are no reptiles, mammals or amphibians on the island, although there are many insect and plant species.
The evolution of two flightless bird species on the island indicates