The correct option is C Low CO2 concentration, accumulation of organic acids resulting in the increased concentration of cell sap.
Scotoactive stomata are the stomata, which open in dark and close during the day time. Scotoactive stomata occur in succulents, e.g., cactus and bryophyllum. Succulents perform crassulacean acid metabolism or CAM in their photosynthesis. Nishida explained, that in succulents during night there is accumulation of organic acids, whereas, during day time the organic acids breakdown releasing carbon dioxide, which keeps stomata closed. During night, because of formation of organic acids carbon dioxide is consumed and concentration of carbon dioxide is less.